1. Australian Antarctic Division Kingston TAS Australia
2. Inst. for Marine and Antarctic Studies, Univ. of Tasmania Hobart TAS Australia
3. Antarctic Climate and Ecosystems Co‐operative Research Centre Hobart TAS Australia
4. Dept of Biological Sciences Macquarie Univ. Sydney NSW Australia
5. Centre d'Etudes Biologiques de Chizé, CNRS Villiers en Bois France