1. Guangdong Provincial Key Laboratory for Plant Epigenetics, Longhua Bioindustry and Innovation Research Institute, College of Life Sciences and Oceanography Shenzhen University Shenzhen 518060 China
2. Key Laboratory of Optoelectronic Devices and Systems of Ministry of Education and Guangdong Province, College of Optoelectronic Engineering Shenzhen University Shenzhen 518060 China
3. Department of Botany and Plant Sciences Institute of Integrative Genome Biology, University of California Riverside 92521 CA USA
4. Department of Biological Sciences, Life Science and Technology Institute Michigan Technological University Houghton 49931 MI USA