1. State Key Laboratory of Microbial Resources and Environmental Microbiology Research Center, Institute of Microbiology; Chinese Academy of Sciences; Beijing 100101 China
2. University of Chinese Academy of Sciences; Beijing 100049 China
3. IMCAS-RCEECAS Joint Laboratory for Environmental Microbial Technology; Beijing China
4. Department of Microbiology and Molecular Genetics; University of California; Davis CA 95616 USA