1. Minnesota Cooperative Fish and Wildlife Research Unit; Department of Fisheries, Wildlife and Conservation Biology; University of Minnesota; St. Paul MN 55108 USA
2. U.S. Fish and Wildlife Service; Fairbanks AK 99701 USA
3. U.S. Geological Survey; Minnesota Cooperative Fish and Wildlife Research Unit; St. Paul MN 55108 USA
4. Eastern Oregon Agriculture & Natural Resource Program; Department of Fisheries and Wildlife; Oregon State University; Union OR 97883 USA
5. Bureau of Land Management; Fairbanks AK 99709 USA