1. Key Laboratory of the Ministry of Education for Coastal and Wetland Ecosystems, College of the Environment and EcologyXiamen University Xiamen Fujian China
2. Third Institute of OceanographyMinistry of Natural Resources Xiamen Fujian China
3. Guangxi Beihai Monitoring and Experimental Station of Marine Ecosystems, Third Institute of OceanographyMinistry of Natural Resources Beihai Guangxi China