1. Department of Genetics; Faculty of Science; Kasetsart University; 50 Phaholyothin Road, Ladyao, Chattuchak, Bangkok 10900 Thailand
2. Royal Forest Department; Sukhothai Province Forest Products Research Center; Sawankhalok, Sukhothai 14110 Thailand
3. Faculty of Science and Technology; Rajamangala University of Technology; 43 Bangpra, Sriracha Chonburi 20110 Thailand
4. School of Biological Science; The University of Queensland, Brisbane St Lucia; QLD 4072 Australia