1. Program in Plant Ecology and Evolution; Department of Ecology and Genetics; Evolutionary Biology Centre; Uppsala University; Norbyvägen 18D 75236 Uppsala Sweden
2. Department of Forest Genetics; Forestry and Forest Products Research Institute (FFPRI); Matsunosato 1 Tsukuba Ibaraki 305-8687 Japan
3. Department of Plant Ecology; Forestry and Forest Products Research Institute (FFPRI); Matsunosato 1 Tsukubaa Ibaraki 305-8687 Japan
4. Laboratory of Forest Ecosystem Studies; Department of Ecosystem Studies; Graduate School of Agriculture and Life Sciences; The University of Tokyo; Yayoi 1-1-1 Bunkyo-ku Tokyo 113-8657 Japan