1. Centre for Sustainability (CSAFE); University of Otago; P.O. Box 56 Dunedin 9054 New Zealand
2. Landcare Research Manaaki Whenua; Private Bag 1930 Dunedin 9054 New Zealand
3. Botany Department; University of Otago; P.O. Box 56 Dunedin 9054 New Zealand