1. NTNU University Museum, Norwegian University of Science and Technology Trondheim NO‐7491 Norway
2. Ecological Genetics Research Unit, Organismal and Evolutionary Biology Research Program, Faculty of Biological and Environmental Sciences University of Helsinki Viikinkaari 1 Helsinki 00790 Finland
3. Natural History Museum University of Oslo Oslo NO‐0318 Norway
4. Department of Systematic and Evolutionary Botany University of Zurich Zurich 8008 Switzerland