1. Ministry of Education Key Laboratory for Biodiversity Science and Ecological Engineering, National Observations and Research Station for Wetland Ecosystems of the Yangtze Estuary, Institute of Eco‐Chongming, School of Life Sciences Fudan University Shanghai 200433 China
2. Key Laboratory of Animal Ecology and Conservation Biology, Institute of Zoology Chinese Academy of Sciences Beijing 100101 China
3. School of Biological Sciences University of Queensland St Lucia Queensland 4072 Australia
4. Current address: Micha V. Jackson, School of Biological Sciences University of Adelaide Adelaide 5005 Australia
5. Global Flyway Network Wakefield 7096 New Zealand