1. Germplasm Bank of Wild Species, Kunming Institute of Botany, Chinese Academy of Science Kunming Yunnan 650201 China
2. Key Laboratory of Medicinal Plant Biology of Yunnan Province Yunnan Agricultural University Kunming Yunnan 650201 China
3. National & Local Joint Engineering Research Center on Germplasm Utilization & Innovation of Chinese Medicinal Materials in Southwestern China Kunming Yunnan 650201 China
4. The Life Science and Technology College, Honghe University Mengzi Yunnan 661199 China
5. Department of Plant Sciences University of Tennessee Knoxville Tennessee 37996 USA