1. Department of Natural History Sciences; Faculty of Science; Hokkaido University; Sapporo; 060-0810; Japan
2. Graduate School of Environmental Earth Science; Hokkaido University; Sapporo; 060-0810; Japan
3. Seian University of Art and Design; Otsu; 520-0248; Japan
4. Zoological Institute; Russian Academy of Sciences; St. Petersburg; 199034; Russia
5. National Institute of Biological Resources; Environmental Research Complex; Incheon; 404-708; South Korea
6. Department of Life Science; Tunghai University; Taichung; 407; Taiwan
7. Laboratory of Wildlife Ecology; Obihiro University of Agriculture and Veterinary Medicine; Obihiro; 080-8555; Japan