1. Institute of Deep-sea Science and Engineering; Chinese Academy of Sciences; Sanya China
2. State Key Laboratory of Freshwater Ecology and Biotechnology; Institute of Hydrobiology Chinese Academy of Sciences; Wuhan China
3. CNRS-University Paul Sabatier; Toulouse Cedex France
4. Freshwater Fisheries Research Center of Chinese Academy of Fishery Sciences; Wuxi China
5. Heilongjiang River Fishery Research Institute; Chinese Academy of Fishery Sciences; Harbin China
6. Institut de Recherche pour le Développement; Unité Mixte de Recherche Biologie des Organismes et Écosystèmes Aquatiques; Muséum National d'Histoire Naturelle; Paris Cedex France