1. Centre for Applied Biological and Social Sciences; State University of Paraiba; Campus V, Horacio Trajano Street, Cristo Redentor 58070-450 João Pessoa Paraiba Brazil
2. MARE-Marine and Environmental Sciences Centre, Faculty of Sciences and Technology; University of Coimbra; 3004-517 Coimbra Portugal
3. Postgraduate Program in Ecology and Conservation; Paraiba State University; 58429-500 Campina Grande Brazil
4. Biology Department & CESAM; University of Aveiro; Campus de Santiago 3810-193 Aveiro Portugal
5. Department of Systematics and Ecology; Federal University of Paraiba; Cidade Universitaria, Castelo Branco 58051-900 João Pessoa Paraiba Brazil