1. School of Life Sciences; University of Warwick; Coventry CV4 7AL UK
2. Université Clermont Auvergne; Université Blaise Pascal; Laboratoire ‘Microorganismes : Génome et Environnement’; BP 10448 Clermont-Ferrand F-63000 France
3. CNRS; UMR 6023; LMGE; Aubière F-63171 France
4. Department of Chemistry and Biomolecular Sciences; Macquarie University; North Ryde NSW 2109 Australia
5. National Oceanography Centre; Hampshire SO14 3ZH UK