1. School of Integrative Plant Science Cornell University Ithaca NY 14853 USA
2. Rice Research Institute & Guangdong Key Laboratory of New Technology in Rice Breeding Guangdong Academy of Agricultural Sciences Guangzhou 510640 China
3. Department of Plant Pathology, College of Plant Protection South China Agricultural University Guangzhou 510642 China
4. Robert W. Holley Center for Agriculture and Health US Department of Agriculture, Agricultural Research Service Ithaca NY 14853 USA