1. University of Tunis El Manar; Faculty of Sciences of Tunis; UR11ES11 Bio-ecology and evolutionary systematic; Tunis Tunisia
2. Regional Activity Centre for Specially Protected Areas (RAC/SPA); Boulevard du Leader Yasser Arafat; Tunis Tunisia
3. University de la Manouba; Higher Institute of Biotechnology of Sidi Thabet; Sidi Thabet; Ariana Tunisia