1. College of Life and Geographic Sciences, Key Laboratory of Biological Resources and Ecology of Pamirs Plateau in Xinjiang Uygur Autonomous Region Kashi University Kashi 844000 China
2. State Key Laboratory of Earth Surface Processes and Resource Ecology, Ministry of Education Key Laboratory for Biodiversity Science and Ecological Engineering, College of Life Sciences Beijing Normal University Beijing 100875 China
3. Yunnan Key Laboratory of Plant Reproductive Adaption and Evolutionary Ecology Yunnan University Kunming 650091 China