1. Department of Entomology and Wildlife Ecology; University of Delaware; Newark DE 19716 USA
2. National Wildlife Research Centre; Environment and Climate Change Canada; Ottawa ON K1A 0H3 Canada
3. Theoretical and Computational Ecology; Institute for Biodiversity and Ecosystem Dynamics; University of Amsterdam; NL-1090 GE Amsterdam the Netherlands
4. U.S. Geological Survey; Patuxent Wildlife Research Center; Laurel MD 20708 USA
5. Department of Biological Sciences; Old Dominion University; Norfolk VA 23529 USA