1. Department of Integrative Brain Science; Graduate School of Medicine; Kyoto University; Konoe-cho, Yoshida, Kyoto-shi Kyoto 606-8501 Japan
2. Center for the Promotion of Interdisciplinary Education and Research; Research and Educational Unit of Leaders for Integrated Medical System; Kyoto University; Kyoto Japan
3. Department of Physiology; Systems Neuroscience Laboratory; Graduate School of Medicine; Hokkaido University; Hokkaido Japan