1. Department of Ecology and Key Laboratory for Earth Surface Processes of the Ministry of Education; College of Urban and Environmental Sciences; Peking University; Beijing 100871 China
2. Center for Macroecology, Evolution and Climate; Natural History Museum of Denmark; University of Copenhagen; Copenhagen Ø DK-2100 Denmark
3. Imperial College London; Grand Challenges in Ecosystems & the Environment Initiative; Silwood Park Campus Ascot Berkshire SL5 7PY UK