1. Department of Fish, Wildlife, and Conservation Biology; Colorado State University; 1474 Campus Delivery Fort Collins CO 80523 USA
2. Proteus Wildlife Research Consultants; P.O. Box 5193 Dunedin 9058 New Zealand
3. U.S. Geological Survey; Patuxent Wildlife Research Center; 12100 Beech Forest Rd Laurel MD 20708 USA