1. Department of Fish, Wildlife, and Conservation Biology Colorado State University Fort Collins CO 80523‐1484 USA
2. U.S. Geological Survey Colorado Cooperative Fish and Wildlife Research Unit Colorado State University Fort Collins CO 80523‐1484 USA
3. Department of Statistics Colorado State University Fort Collins CO 80523‐1484 USA
4. Graduate Degree Program in Ecology Colorado State University Fort Collins CO 80523‐1484 USA
5. Colorado Parks and Wildlife Fort Collins CO 80526 USA
6. National Park Service Fort Collins CO 80525 USA