1. Instituto Gulbenkian de Ciência Oeiras Portugal
2. Current address: Atlantic Technical University (UTA) Mindelo, São Vicente island, Cabo Verde
3. Current address: Department of Evolutionary Biology and Environmental Studies University of Zurich Switzerland
4. UMR 5174 ‐ CNRS, Evolution et Diversité Biologique University Paul Sabatier Toulouse France
5. CE3C: Centre for Ecology, Evolution, and Environmental Changes, Faculty of Sciences University of Lisbon Portugal