1. Animal Demography Unit; Department of Biological Sciences; University of Cape Town; Rondebosch 7701 South Africa
2. Laboratoire ECOMAR; Université de La Réunion; Saint Denis La Réunion
3. Department of Zoology and Entomology; Rhodes University; Grahamstown South Africa
4. DST-NRF Centre of Excellence at the Percy Fitzpatrick Institute of African Ornithology; University of Cape Town; Rondebosch
5. Department of Botany; Rhodes University; Grahamstown South Africa
6. Centre d'Etudes Biologiques de Chizé; Villiers-en-Bois France