1. Animal Ecology/Department of Ecology and Genetics, Evolutionary Biology Centre; Uppsala University; Norbyvägen 18D SE-75236 Uppsala Sweden
2. Department of Evolutionary Ecology; Estación Biológica de Doñana, CSIC; Avda. Américo Vespucio s/n 41092 Sevilla Spain
3. Grupo de Investigación BIOCOST; University of Coruña; 15071 Coruña Spain
4. Current Address: Population Genetics; Institute of Marine Research; 5817 Bergen Norway