1. Key Laboratory of Alpine Ecology and Biodiversity Institute of Tibetan Plateau Research Chinese Academy of Sciences Beijing China
2. Département des Sciences Fondamentales Université du Quebec à Chicoutimi Chicoutimi QC Canada
3. College of Life Sciences Anhui Normal University Wuhu China
4. Key Laboratory of Vegetation Restoration and Management of Degraded Ecosystems Guangdong Provincial Key Laboratory of Applied Botany South China Botanical Garden Chinese Academy of Sciences Guangzhou China
5. CAS Centre for Excellence in Tibetan Plateau Earth Sciences Beijing China
6. Direction de la Recherche Forestière Ministère des Forêts, de la Faune et des Parcs du Québec Québec QC Canada