1. College of Fishery; Guangdong Ocean University; Zhanjiang 524088 People's Republic of China
2. Guangdong Provincial Key Laboratory of Pathogenic Biology and Epidemiology for Aquatic Economic Animals; Zhanjiang 524088 People's Republic of China
3. Guangdong Key Laboratory of Control for Diseases of Aquatic Economic Animals; Zhanjiang 524088 People's Republic of China