1. Department of Production and Population Health; Royal Veterinary College; North Mymms Hatfield Hertfordshire UK
2. Department of Pathology and Pathogen Biology; Royal Veterinary College; North Mymms Hatfield Hertfordshire UK
3. Department of Veterinary Medicine; University of Cambridge; UK
4. Department of Epidemiology and Disease Surveillance; Animal Health Trust; Kentford Newmarket Suffolk UK