1. Department of Veterinary Integrative Biosciences College of Veterinary Medicine and Biomedical Research Texas A&M University College Station TX 77843 USA
2. Department of Population Health and Reproduction School of Veterinary Medicine University of California—Davis Davis CA 95616 USA
3. School of Veterinary Medicine Veterinary Genetics LaboratoryUniversity of California—Davis Davis CA 95616 USA
4. Department of Animal Science University of Nebraska Lincoln NE 68583‐0908 USA