1. Graduate School of Environment and Information Sciences Yokohama National University Yokohama Kanagawa Japan
2. Field Science Center, Graduate School of Agricultural Science Tohoku University Sendai Miyagi Japan
3. Graduate School of Life Sciences Tohoku University Aoba Sendai Miyagi Japan
4. Department of Biological Environment, Faculty of Bioresource Sciences Akita Prefectural University Akita Japan
5. German Centre for Integrative Biodiversity Research (iDiv) Halle‐Jena‐Leipzig Leipzig Germany
6. Institute of Biology, Leipzig University Leipzig Germany