1. School of Geography and the Environment University of Oxford Oxford UK
2. Ecology and Evolutionary Biology Department Yale University New Haven CT USA
3. Biological and Environmental Sciences Faculty of Natural Sciences University of Stirling Stirling UK
4. Institut de Recherche en Ecologie Tropicale CENAREST Libreville Gabon
5. Agence Nationale des Parcs Nationaux Libreville Gabon
6. Biological Sciences Department University of Cape Town Cape Town South Africa