1. Department of Biology Aarhus University Aarhus C Denmark
2. College of Landscape Architecture and Forestry Qingdao Agricultural University Qingdao P.R. China
3. Institute of Ecology and Biodiversity College of Life Sciences Shandong University Qingdao P.R. China
4. Key Laboratory of Coastal Wetland Biogeosciences China Geological Survey Qingdao Institute of Marine GeologyCGSMinistry of Natural Resources Qingdao P.R. China