1. School of Ecological and Environmental Sciences; East China Normal University; Shanghai 200062 China
2. Institute of Agricultural Engineering; Jiangsu University; Zhenjiang Jiangsu 212013 China
3. Center for Global Change and Ecological Forecasting; East China Normal University; Shanghai 200062 China
4. Coastal Ecosystems Research Station of Yangtze River Estuary; Ministry of Education Key Laboratory for Biodiversity Science and Ecological Engineering; Institute of Biodiversity Science; Fudan University; 220 Handan Road Shanghai 200433 China
5. Faculty of Science, Health, Education and Engineering; University of the Sunshine Coast; Maroochydore, DC Qld 4558 Australia