1. Department of Life Sciences Korea University 145 Anamro, Sungbuk‐gu Seoul 02841 Korea
2. National Institute of Agricultural Sciences, Rural Development Administration Jeonju 54874 Korea
3. Department of Horticulture Michigan State University East Lansing MI 48824 USA
4. Department of Plant & Environmental New Resources, College of Life Science and Graduate School of Biotechnology Kyung Hee University Yongin‐si Gyeonggi‐do 17104 Korea
5. Department of Forestry Michigan State University East Lansing MI 48824 USA