1. CREAF, Cerdanyola del Vallès Barcelona 08193 Spain
2. Research School of Biology The Australian National University Canberra 2601 ACT Australia
3. Centre for Tropical Environmental and Sustainability Science, College of Science and Engineering James Cook University Cairns 4878 Qld Australia
4. CSIRO Land and Water Atherton 4883 Qld Australia
5. Geography, Faculty of Environment Science and Economy University of Exeter Laver Building Exeter EX4 4QE UK
6. Departamento de Biologia Vegetal Instituto de Biologia, Universidade Estadual de Campinas (UNICAMP) Campinas 13083‐970 SP Brazil
7. School of Geosciences University of Edinburgh Edinburgh EH9 3FF UK