1. Department of Biological and Environmental Science; Centre of Excellence in Biological Interactions; University of Jyväskylä; Jyväskylä Finland
2. Centre d'Ecologie et de Sciences de la Conservation UMR 7204 CNRS/MNHN/UPMC; Muséum National d'Histoire Naturelle; Paris France
3. Biogéosciences; UMR CNRS 6282; Université de Bourgogne; Dijon France
4. Department of Ecology; Swedish University of Agricultural Sciences; Uppsala Sweden