1. Department of Behavioural Ecology and Evolutionary Genetics; Max Planck Institute for Ornithology; Eberhard-Gwinner-Str. 82319 Seewiesen Germany
2. Department of Evolutionary Biology; Bielefeld University; Morgenbreede 45 33615 Bielefeld Germany
3. Department of Evolutionary Biology; Uppsala University; Norbyvägen 18D 752 36 Uppsala Sweden