1. Institute of Evolution & Marine Biodiversity, and College of Fisheries Ocean University of China Qingdao 266003 China
2. School of Life Sciences East China Normal University Shanghai 200241 China
3. Department of Marine Organism Taxonomy and Phylogeny Institute of Oceanology Chinese Academy of Sciences Qingdao 266071 China
4. Marine College Shandong University Weihai 264209 China
5. Department of Life Sciences Natural History Museum London SW7 5BD United Kingdom