1. South African Institute for Aquatic Biodiversity (SAIAB); Private Bag 1015, Grahamstown, 6140 South Africa
2. Department of Zoology & Entomology; University of Fort Hare; Private Bag X1314, Alice, 5700 South Africa
3. South African Environmental Observation Network (SAEON); Elwandle Coastal Node, P.O. Box 77 000, Port Elizabeth, 6031 South Africa
4. School of Science, Monash University Malaysia; 47500, Bandar Sunway Selangor Darul Ehsan Malaysia
5. Department of Zoology; Institute for Coastal and Marine Research (CMR), Nelson Mandela University; P.O. Box 77 000, Port Elizabeth, 6031 South Africa