1. National Institute of Polar Research 10‐3 Midori‐cho, Tachikawa, Tokyo 190‐8518 Japan
2. Department of Ecology and Evolutionary Biology University of California Santa Cruz, Long Marine Lab 130 McAllister Way, Santa Cruz, California 95060 U.S.A.
3. Department of Medicine University of California San Diego, 9500 Gilman Drive, La Jolla, California 92093 U.S.A.