1. The Key Laboratory of Biomedical Information Engineering, Ministry of Education, School of Life Science and Technology; Xi'an Jiaotong University; Xi'an 710049 China
2. University of Innsbruck, Research Institute for Limnology; Mondseestr. 9 Mondsee 5310 Austria
3. Institute of Evolution & Marine Biodiversity; Ocean University of China; Qingdao 266003 China
4. Key Laboratory of Aquatic Biodiversity and Conservation of Chinese Academy of Sciences, Institute of Hydrobiology; Chinese Academy of Sciences; Wuhan 430072 China
5. Zoology Department, College of Science; King Saud University; P. O. Box 2455 Riyadh 11451 Saudi Arabia