1. Department of Bioscience; Aarhus University; Frederiksborgvej 399 DK-4000 Roskilde Denmark
2. Helmholtz Centre for Environmental Research - UFZ; Department of Ecological Modelling; Permoserstraße 15 04318 Leipzig Germany
3. School of Biological Sciences, University of Reading, Harborne Building; University of Reading; Whiteknights Reading Berkshire, RG6 6AS United Kingdom
4. Lighthouse Field Station, Institute of Biological and Environmental Sciences; University of Aberdeen; Cromarty IV11 8YL United Kingdom