1. Graduate School of Agriculture Science; Tohoku University; 1-1 Tsutsumidori-Amamiyamachi Aoba-ku Sendai 981-8555 Japan
2. RIKEN Center for Sustainable Resource Science; 1-7-22 Suehiro-cho Tsurumi-ku Yokohama 230-0045 Japan
3. Faculty of Life and Environmental Sciences; Tsukuba University; 1-1-1 Tenodai Tsukuba 305-8572 Japan
4. Japan Science and Technology Agency; Precursory Research for Embryonic Science and Technology (PRESTO); Kawaguchi Saitama 332-0012 Japan
5. Graduate School of Life Sciences; Tohoku University; Katahira 2-1-1 Aoba-ku Sendai 980-8577 Japan
6. Graduate School of Agricultural and Life Sciences; University of Tokyo; Bunkyo-ku Tokyo 113-8652 Japan