1. AMAP Université Montpellier CIRAD CNRS INRAE IRDUMR AMAP Boulevard de La Lironde34098Montpellier France
2. Botany Department School of Natural Sciences Trinity College Dublin Dublin 2 Ireland
3. Division of Paleobotany, Biodiversity Institute University of Kansas Lawrence KS 66047 USA
4. Department of Ecology and Evolutionary Biology Natural History Museum and Biodiversity Institute University of Kansas Lawrence KS 66047 USA