1. Molecular Ecology and Evolution Programme; Department of Genetics; University of Pretoria; Pretoria South Africa
2. Université de La Réunion; Laboratoire ECOMAR - FRE CNRS 3560; Labex CORAIL; CS 92003 97744 Saint-Denis La Réunion France
3. USR3278-CRIOBE-CNRS-EPHE; Laboratoire d'excellence ‘CORAIL’; Université de Perpignan-CBETM; 66860 Perpignan Cedex France
4. Florida Museum of Natural History; Gainesville FL 32611-7800 USA