1. UMR Biologie des Organismes et des Ecosystèmes Aquatiques (MNHN-IRD-CNRS-UPMC); Muséum National d’Histoire Naturelle; 43 rue Cuvier-CP 26 75005 Paris France
2. North Carolina State Museum of Natural Sciences; 11 West Jones St Raleigh NC 27601 U.S.A.
3. Department of Ecology and Evolutionary Biology; Princeton University; Princeton NJ 08544 U.S.A.
4. Raffles Museum of Biodiversity Research; National University of Singapore; S117543 Singapore