1. Key Laboratory of Exploration and Utilization of Aquatic Genetic Resources; Shanghai Ocean University, Ministry of Education Shanghai; Shanghai 201306 China
2. Department of Biological Sciences; Louisiana State University; LA 70803 USA
3. Institute of Botany; the Chinese Academy of Sciences; Beijing 100093 China
4. National Key Laboratory of Plant Molecular Genetics, Institute of Plant Physiology and Ecology; Shanghai Institutes for Biological Sciences, the Chinese Academy of Sciences; Shanghai 200032 China