1. Key Laboratory of Plant Resources Conservation and Sustainable Utilization; South China Botanical Garden, Chinese Academy of Sciences; Guangzhou 510650 P. R. China
2. Key Laboratory of Plant Germplasm Enhancement and Specialty Agriculture; Wuhan Botanical Garden, Chinese Academy of Sciences; Wuhan Hubei 430074 P. R. China
3. National Key Laboratory of Biochemical Engineering; Institute of Process Engineering, Chinese Academy of Sciences; Beijing 100190 P. R. China