1. EPHE PSL Research University UPVD CNRS USR 3278 CRIOBE BP 1013 98729 Papetoai Moorea French Polynesia
2. Laboratoire d'excellence “CORAIL” EPHE PSL Research University UPVD CNRS USR 3278 CRIOBE, BP 1013 98729 Papetoai, Moorea French Polynesia
3. MARE – Marine and Environmental Sciences Centre Faculdade de Ciências da Universidade de Lisboa Campo Grande, Lisboa 1749‐016 Portugal
4. Coastal Research Center Department of Ecology, Evolution and Marine Biology Marine Science Institute University of California Santa Barbara Santa Barbara CA 93106 USA
5. Department of Ecology and Evolutionary Biology University of California Santa Cruz 100 Shaffer Road Santa Cruz CA 95060 USA